Abstract: Observations on the developmental process of the e female and male gametophytes in Astragalus membranaceus were carried but by the optical microscope, in the meantime, made cytochemical experiment for the pollen development, the main results are as follows:1. At first mitosis of pollen, a smaller hemispherical generative cell and a larger vegetative cell were formed. 2. During generative cell development, structure of cell wall was temporary. that the wall was tested by PAS reaction and by the fluorescence reaction with aniline blue always showed negative, namely, generative cell wall didn't show the properties of celluse or callose wall. 3. Course of the disintegration of wall between vegetative cell and generative cell aswell as that generative cell came into the vegetative cell. 4. In the maturepollen, generative nucleus give a strong positive Feulgen reaction, but vegetative nucleus give a weak positive Feulgen reaction. 5. Development of female gametophytes originate from subhypoclermal archesporial cells oriented in the nucellus, but only one of these directly develops into the Polyonum type embryo sac. The retativity of the external morphology of the flower bud and the development of the female and male gametophytes in Astragalus membranoccus were compared.