植物研究 ›› 2025, Vol. 45 ›› Issue (2): 254-265.doi: 10.7525/j.issn.1673-5102.2025.02.011
• 研究论文 • 上一篇
曾艳丽1, 关艳辉2, 乔璐靖1, 陈雅婷1, 琚存勇1(), 蔡体久1
收稿日期:
2024-09-16
出版日期:
2025-03-20
发布日期:
2025-04-04
通讯作者:
琚存勇
E-mail:qucy09@nefu.edu.cn
作者简介:
曾艳丽(1998—),女,硕士研究生,主要从事森林植物资源保护与利用研究。
基金资助:
Yanli ZENG1, Yanhui GUAN2, Lujing QIAO1, Yating CHEN1, Cunyong JU1(), Tijiu CAI1
Received:
2024-09-16
Online:
2025-03-20
Published:
2025-04-04
Contact:
Cunyong JU
E-mail:qucy09@nefu.edu.cn
摘要:
研究林木径向生长与其近邻体结构的关系,有助于了解林分结构的细微之处,使森林经营更加定量化和精确化。以小北湖自然保护区内黄花落叶松(Larix olgensis)林和蒙古栎(Quercus mongolica)林为研究对象,计算优势、伴生树种不同径级林木的径向生长量与近邻体结构参数,采用通径分析探究近邻体结构对不同径级林木径向生长的直接与间接影响。结果表明:无论是作为优势树种还是伴生树种,黄花落叶松和蒙古栎的小径级林木径向生长量均要低于中、大径级,其中,黄花落叶松的小径级林木径向生长量要低于蒙古栎,中、大径级则要高于蒙古栎。小、中径级林木径向生长与近邻体结构显著相关,大径级林木则不相关。开阔比数和竞争指数是影响黄花落叶松和蒙古栎径向生长的主要因子,且对伴生树种的影响程度要高于优势树种。随着径级增大,近邻体结构对径向生长的影响程度逐渐减小。此外,开阔比数与竞争指数的间接相互作用也会影响黄花落叶松与蒙古栎的径向生长。因此,该地区进行森林经营时,应选择小、中径级林木进行近邻体结构调整,以提高它们的开阔比数和降低竞争指数为主。
中图分类号:
曾艳丽, 关艳辉, 乔璐靖, 陈雅婷, 琚存勇, 蔡体久. 近邻体结构对不同径级黄花落叶松和蒙古栎径向生长的影响[J]. 植物研究, 2025, 45(2): 254-265.
Yanli ZENG, Yanhui GUAN, Lujing QIAO, Yating CHEN, Cunyong JU, Tijiu CAI. Effects of Neighborhood Structure on Radial Growth of Larix olgensis and Quercus mongolica with Different Diameter Classes[J]. Bulletin of Botanical Research, 2025, 45(2): 254-265.
表1
样地基本信息
样地类型 Sample plot type | 北纬 North latitude | 东经 East longitude | 郁闭度 Canopy density | 密度 Density/(tree·hm-2) | 平均胸径 Mean DBH/cm | 平均树高Mean tree height/m | 树种组成 Tree species composition |
---|---|---|---|---|---|---|---|
黄花落叶松林 L. olgensis forest | 44°9′51.55″ | 128°35′50.24″ | 0.7 | 876 | 17.90 | 14.00 | 9落1蒙 9 Lo 1 Qm |
蒙古栎林 Q. mongolica forest | 44°10′44.35″ | 128°37′6.82″ | 0.6 | 879 | 17.00 | 9.20 | 9蒙1落+黑 9 Qm1 Lo+Bd |
表2
不同径级优势、伴生黄花落叶松与蒙古栎近邻体结构和径向生长量的通径分析
径级 Diameter class | 近邻体结构指数 Neighborhood structural index | 优势黄花落叶松 DLo | 伴生黄花落叶松 ALo | 优势蒙古栎 DQm | 伴生蒙古栎 AQm | ||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
直接路径系数 Direct path coefficient | 间接路径系数 Indirect path coefficient | 直接路径系数 Direct path coefficient | 间接路径系数 Indirect path coefficient | 直接路径系数 Direct path coefficient | 间接路径系数 Indirect path coefficient | 直接路径系数 Direct path coefficient | 间接路径系数 Indirect path coefficient | ||||||||||||||
CI | Msi | W | OP | CI | Msi | W | OP | CI | Msi | W | OP | CI | Msi | W | OP | ||||||
小径级 Small diameter class | CI | -0.183 | -0.007 | -0.029 | 0.079 | -0.454 | -0.093 | 0.053 | 0.268 | -0.216 | -0.008 | -0.011 | 0.082 | -0.446 | -0.094 | -0.100 | 0.052 | ||||
Msi | -0.162 | -0.006 | -0.018 | -0.059 | 0.180 | 0.037 | -0.013 | -0.042 | 0.016 | 0.001 | -0.002 | 0 | -0.077 | -0.016 | -0.011 | 0.020 | |||||
W | -0.202 | -0.032 | -0.022 | -0.008 | 0.031 | -0.004 | -0.002 | -0.003 | -0.060 | -0.003 | 0.008 | 0.005 | 0.222 | 0.050 | 0.030 | -0.021 | |||||
OP | 0.027 | -0.012 | 0.010 | 0.001 | 0.084 | -0.050 | -0.019 | -0.007 | 0.251 | -0.095 | -0.002 | -0.022 | 0.594 | -0.069 | -0.151 | -0.056 | |||||
中径级 Medium diameter class | CI | -0.186 | 0.016 | 0.008 | -0.174 | 0.048 | 0.068 | 0.016 | -0.127 | 0.010 | 0.014 | 0.037 | -0.079 | -0.021 | 0.002 | 0.008 | |||||
Msi | -0.049 | 0.004 | 0.008 | -0.004 | -0.098 | 0.027 | -0.011 | 0.031 | -0.052 | 0.004 | -0.005 | 0.009 | 0.390 | 0.105 | -0.050 | 0.021 | |||||
W | -0.046 | 0.002 | 0.007 | -0.004 | 0.084 | -0.033 | 0.009 | -0.019 | 0.031 | -0.003 | 0.003 | 0.004 | -0.162 | 0.004 | 0.021 | 0.001 | |||||
OP | 0.282 | -0.147 | 0.024 | 0.022 | 0.308 | -0.028 | -0.096 | -0.071 | 0.243 | -0.070 | -0.039 | 0.035 | 0.390 | -0.041 | 0.021 | -0.003 | |||||
大径级 Large diameter class | CI | 0.135 | -0.050 | 0.007 | 0.001 | 0.081 | -0.022 | 0.030 | -0.045 | 0.006 | 0.002 | 0 | 0 | 0 | 0 | 0 | |||||
Msi | 0.115 | -0.042 | 0.001 | 0.008 | -0.179 | 0.049 | -0.007 | -0.047 | 0.006 | -0.002 | -0.002 | 0 | 0 | 0 | 0 | ||||||
W | 0.063 | 0.003 | 0 | -0.002 | 0.264 | 0.098 | 0.010 | 0.024 | -0.001 | 0.001 | 0.001 | 0 | 0 | 0 | 0 | ||||||
OP | 0.033 | 0 | 0.002 | -0.001 | -0.051 | 0 | -0.002 | -0.001 | 0 | 0 | 0 | 0 |
1 | GSCHWANTNER T, RIEDEL T, HENNING L,et al.Improved large-area forest increment information in Europe through harmonisation of National Forest Inventories[J].Forest Ecology and Management,2024,562:121913. |
2 | 冯婷华,郭晋平,常建国.中条山辽东栎优势木径向生长的时空变异特征[J].山西农业科学,2022,50(4):559-567. |
FENG T H, GUO J P, CHANG J G.Spatial and temporal variation characteristics of radial growth of plus trees of Quercus liaotungensis in Zhongtiao Mountain[J].Journal of Shanxi Agricultural Sciences,2022,50(4):559-567. | |
3 | 池静姚,潘磊磊, SEMYUNG K,等.呼伦贝尔沙地樟子松天然林结构多样性和竞争对树木生长的影响[J].中国沙漠,2024,44(5):29-40. |
CHI J Y, PAN L L, SEMYUNG K,et al.Influence of stand structure diversity and competition on tree growth of natural Pinus sylvestris var.mongolica forest in Hulun Buir Sandy Land[J].Journal of Desert Research,2024,44(5):29-40. | |
4 | WANG Z C, LI Y X, WANG G Y,et al.Drivers of spatial structure in thinned forests[J].Forest Ecosystems,2024,11:100182. |
5 | 蔡礼蓉,匡旭,房帅,等.长白山阔叶红松林3个常见树种径向生长的影响因素[J].应用生态学报,2017,28(5):1407-1413. |
CAI L R, KUANG X, FANG S,et al.Factors influencing tree radial growth of three common species in broad-leaved Korean pine mixed forests in Changbai Mountains,China[J].Chinese Journal of Applied Ecology,2017,28(5):1407-1413. | |
6 | QI J H, FAN Z X, FU P L,et al.Differential determinants of growth rates in subtropical evergreen and deciduous juvenile trees:carbon gain,hydraulics and nutrient-use efficiencies[J].Tree Physiology,2021,41(1):12-23. |
7 | 张萌,范秀华.吉林蛟河针阔混交林红松和紫椴个体生长的影响因素[J].北京林业大学学报,2024,46(9):26-34. |
ZHANG M, FAN X H.Factors affecting individual growth of Pinus koraiensis and Tilia amurensis in coniferous and broadleaved mixed forests in Jiaohe,Jilin Province of northeastern China[J].Journal of Beijing Forestry University,2024,46(9):26-34. | |
8 | 韩大校,金光泽.地形和竞争对典型阔叶红松林不同生长阶段树木胸径生长的影响[J].北京林业大学学报,2017,39(1):9-19. |
HAN D X, JIN G Z.Influences of topography and competition on DBH growth in different growth stages in a typical mixed broadleaved-Korean pine forest,Northeastern China[J].Journal of Beijing Forestry University,2017, 39(1):9-19. | |
9 | 余枭,欧阳勋志,潘萍,等.庐山常绿阔叶林不同生长阶段林木空间结构特征及其评价[J].北京林业大学学报,2022,44(12):32-40. |
YU X, OUYANG X Z, PAN P,et al.Spatial structure characteristics and its evaluation of evergreen broadleaved forest at different growth stages in Lushan Mountain,Jiangxi Province of Eastern China[J].Journal of Beijing Forestry University,2022,44(12):32-40. | |
10 | 谢伊,杨华.长白山天然云冷杉针阔混交林主要树种胸径生长与林分空间结构的关系[J].北京林业大学学报,2022,44(9):1-11. |
XIE Y, YANG H.Relationship between stand spatial structure and DBH increment of principal species in natural spruce-fir mixed forest in Changbai Mountains of Northeastern China[J].Journal of Beijing Forestry University,2022,44(9):1-11. | |
11 | 董雪婷,张静,张志东,等.树种相互作用、林分密度和树木大小对华北落叶松生产力的影响[J].应用生态学报,2021,32(8):2722-2728. |
DONG X T, ZHANG J, ZHANG Z D,et al.Effects of tree species interaction,stand density,and tree size on the productivity of Larix principis-rupprechtii [J].Chinese Journal of Applied Ecology,2021,32(8):2722-2728. | |
12 | 吕延杰,杨华,张青,等.云冷杉天然林林分空间结构对胸径生长量的影响[J].北京林业大学学报,2017,39(9):41-47. |
LÜ Y J, YANG H, ZHANG Q,et al.Effects of spatial structure on DBH increment of natural spruce-fir forest[J].Journal of Beijing Forestry University,2017,39(9):41-47. | |
13 | LIN S, LI Y, CHEN M F,et al.Effects of local neighborhood structure on radial growth of Picea crassifolia Kom.and Betula platyphylla Suk.plantations in the loess alpine region,China[J].Forest Ecology and Management,2021,491:119195. |
14 | ZHANG G Q, HUI G Y, YANG A M,et al.A simple and effective approach to quantitatively characterize structural complexity[J].Scientific Reports,2021,11(1):1326. |
15 | 惠刚盈.基于相邻木关系的林分空间结构参数应用研究[J].北京林业大学学报,2013,35(4):1-8. |
HUI G Y.Studies on the application of stand spatial structure parameters based on the relationship of neighborhood trees[J].Journal of Beijing Forestry University,2013,35(4):1-8. | |
16 | 王立凤,刘丹,陈鑫,等.黑龙江小北湖国家级自然保护区的植物多样性[J].贵州农业科学,2020,48(6):131-134. |
WANG L F, LIU D, CHEN X,et al.Plant diversity of Xiaobeihu National Nature Reserve in Heilongjiang Province[J].Guizhou Agricultural Sciences,2020,48(6):131-134. | |
17 | TANG G R, ZHANG H, XING H L,et al.Light adaptation strategies of Quercus mongolica at different ages in four plantations[J].Forestry Research,2024,4(1):e005. |
18 | 王剑武,徐森,季碧勇,等.地形和林分空间结构对浙江省天然阔叶混交林主要先锋树种胸径生长的影响[J].应用生态学报,2024,35(2):298-306. |
WANG J W, XU S, JI B Y,et al.Effects of topography and stand spatial structure on the diameter at breast height growth of major pioneer tree species of natural broad-leaved mixed forests in Zhejiang Province,China[J].Chinese Journal of Applied Ecology,2024,35(2):298-306. | |
19 | 汤孟平,陈永刚,施拥军,等.基于Voronoi图的群落优势树种种内种间竞争[J].生态学报,2007,27(11):4707-4716. |
TANG M P, CHEN Y G, SHI Y J,et al.Intraspecific and interspecific competition analysis of community dominant plant populations based on Voronoi diagram[J].Acta Ecologica Sinica,2007,27(11):4707-4716. | |
20 | 宋语涵,张晨,蔡体久,等.基于Voronoi图的阔叶红松林空间结构特征分析[J].北京林业大学学报,2021,43(1):20-26. |
SONG Y H, ZHANG C, CAI T J,et al.Quantitative analysis of spatial structural characteristics of broadleaved Korean pine forest based on Voronoi diagram[J].Journal of Beijing Forestry University,2021,43(1):20-26. | |
21 | 于帅,蔡体久,张丕德,等.边缘校正方法对空间结构参数影响的尺度效应[J].林业科学,2023,59(10):57-65. |
YU S, CAI T J, ZHANG P D,et al.Effects of edge correction methods on spatial structure parameters[J].Scientia Silvae Sinicae,2023,59(10):57-65. | |
22 | 张明辉,尹昀洲,王珂,等.水曲柳人工林空间结构特征对土壤养分含量的影响[J].北京林业大学学报,2023,45(9):73-82. |
ZHANG M H, YIN Y Z, WANG K,et al.Effects of spatial structure characteristics of Fraxinus mandshurica plantation on soil nutrient content[J].Journal of Beijing Forestry University,2023,45(9):73-82. | |
23 | 张毅锋,汤孟平.天目山常绿阔叶林空间结构动态变化特征[J].生态学报,2021,41(5):1959-1969. |
ZHANG Y F, TANG M P.Analysis on spatial structure dynamic characteristics of evergreen broad-leaved forest in Tianmu Mountain[J].Acta Ecologica Sinica,2021,41(5):1959-1969. | |
24 | 汤孟平,唐守正,雷相东,等.两种混交度的比较分 析[J].林业资源管理,2004(4):25-27. |
TANG M P, TANG S Z, LEI X D,et al.Comparison analysis on two minglings[J].Forest Resources Management,2004(4):25-27. | |
25 | 惠刚盈, VON GADOW K, ALBERT M.角尺度:一个描述林木个体分布格局的结构参数[J].林业科学,1999,35(1):37-42. |
HUI G Y, VON GADOW K, ALBERT M.The neighbourhood pattern:a new structure parameter for describing distribution of forest tree position[J].Scientia Silvae Sinicae,1999,35(1):37-42. | |
26 | 张泽鑫,蔡有柱,赵丽娟,等.祁连山东部森林林分结构和环境因素对草本物种多样性的影响[J].生态学报,2024,44(5):2089-2099. |
ZHANG Z X, CAI Y Z, ZHAO L J,et al.Effects of stand structure and environmental factors on understory herbaceous plant diversity in eastern Qilian Mountains[J].Acta Ecologica Sinica,2024,44(5):2089-2099. | |
27 | 胡艳波,惠刚盈,王宏翔,等.随机分布的角尺度置信区间及其应用[J].林业科学研究,2014,27(3):302-308. |
HU Y B, HUI G Y, WANG H X,et al.Uniform angle index(W) confidence interval of the random distribution and its application[J].Forest Research,2014,27(3):302-308. | |
28 | 胡艳波.基于结构化森林经营的天然异龄林空间优化经营模型研究[D].北京:中国林业科学研究院,2010. |
HU Y B.Structure-based spatial optimization management model for natural uneven-aged forest[D].Beijing:Chinese Academy of Forestry,2010. | |
29 | 谢旭强,王庆成,李秋雨,等.水曲柳和黄菠萝苗木生长及光合生理对不同光强的响应[J].森林工程,2023,39(3):73-81. |
XIE X Q, WANG Q C, LI Q Y,et al.Response of growth and photosynthetic physiology of Fraxinus mandshurica and Phellodendron amurense seedlings to different light intensities[J].Forest Engineering,2023,39(3):73-81. | |
30 | 徐磊,吴小云,律江,等.散射辐射比例对华北平原杨树人工林生态系统能量分配的影响[J].林业科学,2024,60(3):100-110. |
XU L, WU X Y, LÜ J,et al.Impacts of diffuse radiation fraction on energy partitioning in a poplar plantation in the North China Plain[J].Scientia Silvae Sinicae,2024,60(3):100-110. | |
31 | 解书文,金光泽,刘志理.小兴安岭不同径级白桦枝叶性状变异及权衡[J].北京林业大学学报,2023,45(12):32-40. |
XIE S W, JIN G Z, LIU Z L.Variations and trade-offs of twig-leaf traits for Betula platyphylla with different diameter classes in Xiaoxing’an Mountains of Northeastern China[J].Journal of Beijing Forestry University,2023,45(12):32-40. | |
32 | 刘澳,王嘉铮,卢思航,等.长汀生态恢复区不同混交比例针阔混交林内马尾松种群动态及其驱动机制[J].林业科学,2024,60(5):89-97. |
LIU A, WANG J Z, LU S H,et al.Population dynamics and driving mechanism of Pinus massoniana in coniferous and broad-leaved mixed forests with different mixing ratios in Changting ecological restoration area[J].Scientia Silvae Sinicae,2024,60(5):89-97. | |
33 | ZHANG H T, OVASKAINEN O, CHI X L,et al.Spatiotemporal variation in the negative effect of neighbourhood crowding on stem growth[J].Journal of Ecology,2024,112(5):1140-1149. |
34 | PIPONIOT C, ANDERSON-TEIXEIRA K J, DAVIES S J,et al.Distribution of biomass dynamics in relation to tree size in forests across the world[J].New Phytologist,2022,234(5):1664-1677. |
35 | LIN S, LI Y, LI Y H,et al.Influence of tree size,local forest structure,topography,and soil resource availability on plantation growth in Qinghai Province,China[J].Ecological Indicators,2021,120:106957. |
36 | ZHANG J, ZHAO J M, CHENG R M,et al.Effects of neighborhood competition and stand structure on the productivity of pure and mixed Larix principis-rupprechtii forests[J].Forests,2022,13(8):1318. |
37 | MADRIGAL-GONZÁLEZ J, RUIZ-BENITO P, RATCLIFFE S,et al.Complementarity effects on tree growth are contingent on tree size and climatic conditions across Europe[J].Scientific Reports,2016,6(1):32233. |
38 | SHEN Z Q, LI Y Z, CHEN Z Y,et al.Species identity and initial size rather than neighborhood interactions influence survival in a response-surface examination of competition[J].Frontiers in Plant Science,2020,11:1212. |
39 | ZHANG H N, ZHANG X S, LÜ Y Y,et al.How topography and neighbor shape the fate of trees in subtropical forest restoration:environmental filtering and resource competition drive natural regeneration[J].Forest Ecosystems,2024,11:100169. |
40 | 姚慧芳,卢杰,曾加芹,等.藏东南川滇高山栎天然林的种内与种间竞争指数的海拔差异[J].林业科学,2022,58(8):53-62. |
YAO H F, LU J, ZENG J Q,et al.Altitudinal differences in intraspecific and interspecific competition index of natural forests of Quercus aquifolioides in Southeast Tibet[J].Scientia Silvae Sinicae,2022,58(8):53-62. | |
41 | DE GROOTE S R E, VANHELLEMONT M, BAETEN L,et al.Competition,tree age and size drive the productivity of mixed forests of pedunculate oak,beech and red oak[J].Forest Ecology and Management,2018,430:609-617. |
42 | LARSON A J, LUTZ J A, DONATO D C,et al.Spatial aspects of tree mortality strongly differ between young and old-growth forests[J].Ecology,2015,96(11):2855-2861. |
43 | 刘万生,李想,陈福元,等.蒙古栎林种内和种间竞争研究[J].植物研究,2020,40(4):552-558. |
LIU W S, LI X, CHEN F Y,et al.Intraspecific and interspecific competition of Quercus mongolica forest[J].Bulletin of Botanical Research,2020,40(4):552-558. | |
44 | ZHANG H T, GHEYRET G, BAI Y H,et al.Functional dissimilarity in mixed forests promotes stem radial growth by mitigating tree water deficit[J].National Science Review,2024,11(3):nwad320. |
45 | ZHAN H, LIU H Z, WANG T C,et al.Selection and validation of reference genes for quantitative real-time PCR of Quercus mongolica Fisch.ex Ledeb under abiotic stresses[J].PLoS One,2022,17(4):e0267126. |
46 | 周浩然,孙洪刚,张鹏,等.混交对杉木和山杜英混交林生长、结构和生产力的影响[J].植物研究,2024, 44(2):307-320. |
ZHOU H R, SUN H G, ZHANG P,et al.Effect of species-mixing on the growth,structure and productivity of Cunninghamia lanceolata and Elaeocarpus sylvestris mixed forests[J].Bulletin of Botanical Research,2024,44(2):307-320. | |
47 | COOMES D A, ALLEN R B.Effects of size,competition and altitude on tree growth[J].Journal of Ecology,2007,95(5):1084-1097. |
48 | RUBIO-CUADRADO Á, MONTES F, ALBERDI I,et al.Analyses from stand to tree level allow disentangling the effects of age,size,origin and competition on tree growth sensitivity to climate in natural and afforested Scots pine forests[J].Agricultural and Forest Meteorology,2024,355:110148. |
49 | FORRESTER D I.Linking forest growth with stand structure:tree size inequality,tree growth or resource partitioning and the asymmetry of competition[J].Forest Ecology and Management,2019,447:139-157. |
50 | ALI A.Forest stand structure and functioning:current knowledge and future challenges[J].Ecological Indicators,2019,98:665-677. |
51 | LANG A C, HÄRDTLE W, BARUFFOL M,et al.Mechanisms promoting tree species co-existence:experimental evidence with saplings of subtropical forest ecosystems of China[J].Journal of Vegetation Science,2012,23(5):837-846. |
[1] | 任天梦, 鲍雨, 苏亚勋, 陈庆斌, 魏立颖, 纪若璇, 于笑, 夏新莉, 尹伟伦, 刘超. 不同修枝处理对10年生三倍体毛白杨生长和光合特性的影响[J]. 植物研究, 2023, 43(6): 846-856. |
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